The river flows initially southwards for 1 km (0.6 mi) to a small lake at Baldhorns Park, before running eastwards through a largely rural area towards Crawley.
The first tributaries to join the young river drain the northernmost part of St Leonard's Forest, between Horsham and Crawley, although much of the forest is in the catchment area of the River Arun.
The River Mole is a tributary of the River Thames in southern England.
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From Painshill Park the river flows northeastwards to the Thames, passing to the west of Esher Commons and then forming the boundary between Hersham and Esher, where the mean flow of the river is 5.43 m/s (192 cu ft/s).
In response to heavy flooding of East Molesey and Thames Ditton in September 1968, the river was modified downstream of Albany Bridge to the Thames and new flood defences were constructed.
The meander was reinstated in 1997, in an attempt to enhance this area of the Park, although it has since become blocked by silt.
The entirety of the Mole Gap lies within the Surrey Hills Area of Outstanding Natural Beauty.
The 18th-century weir at Betchworth was modified in 2004 to facilitate the installation of two 27.5 k W low-head hydro turbines.
About 90% of the energy generated is fed into the regional electricity grid, while the remainder is used to supply the Betchworth Park Estate, where the weir is situated.The course of the Mole within the airport perimeter has been altered several times since commercial flights began in 1945; however the meanders visible on the 1839 tithe map in the 1.5 km stretch immediately north of the runway were reinstated in 1999, in a £1.2 million project to facilitate airport expansion.The Mole enters Surrey to the south of Horley, where it meets the Gatwick Stream, a tributary draining Worth Forest to the southeast of Crawley.There is only one aquifer in the drainage basin, at Fetcham, which means that the majority of the water in the river is from surface drainage, particularly from Gatwick Airport and the urban areas of Horley and Crawley, and that the flow rate responds rapidly to rainfall.River Mole rises in Baldhorns Copse 0.7 km (0.4 mi) to the south of the village of Rusper in West Sussex.Water is able to flow out of the river through swallow holes in the bed and banks, decreasing the volume of water carried in the main channel.